
From left to right: Lioness 3,4,6 and 7
These were probably intended to be lionesses, but I see nothing feminine about them. Perhaps they are maneless lions that helped Zira fend off rogue male lions since Nuka was too weak and Kovu was too young? Maybe they're related to Zira? Her sons, but like Nuka hey were unhealthy thus lost(or were born without) their manes. Maybe her brothers? She was their older sister and since Sanaa(my version of Zira's mom) was dead and Amri(my version of Zira's dad) had been Driven off for killling her she raised them through Scar's reign and once they were seen as a threat they were exiled. Once Simba came back, he exiles Zira and she finds her brother's in the outlands. They have started a pride of their own but have no real leadership skills so apoint her as the leader since she raised them and was once Scar's queen.
This could be debated over as their chest fur extends to their muzzles but males have been known to have this. Example: Tojo. Their whiskers are gone but some males in the TLK unvirse have been seen with all the male traits except whiskers. (See the comic: An unusual choir)
Also, they have nose bridges similar to Nuka's. One of them may have fathered Nuka? And they are bigger than the lionesses around them. Their eyeshades are dark and Scar shaped. Maybe Zira's earlier sons from Scar?